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Paper published on the modelling of counter-shading camouflage

We have had a paper published (online ahead-of-print) in the American Naturalist, on what makes counter-shading camouflage optimal:

Three-Dimensional Camouflage: Exploiting Photons to Conceal Form Olivier Penacchio, P. George Lovell, Innes C. Cuthill, Graeme D. Ruxton, and Julie M. Harris